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r is the relatedness of the recipient to the actor (what proportion of genes they share) - a sister, parent or offsping (for a diploid species) would have an r of 1/2, a cousin would have an r of 1/8, an identical twin or a clone would have an r of 1 and so on. (r can never be greater than 1 since you can't be more related to someone else than you are to yourself - thus "devalued")
Therefore, if the recipient in an individual case were the sister of the actor then the benefit would have to be double the cost to the actor. In the case of cousins the benefit would have to be eight times the cost to the actor and so on.
Sister: 1/2B > C Cousin: 1/8B > C
Permalink Updated: Tue, 31 Aug 2010 05:39:27 UTC | #508364